As a proton moves in the direction of the electric field lines, it is moving from low to high potential and gains electric potential energy.
The imaginary lines are of an electric field such that the tangent drawn to it points in the direction of electrostatic force.
When a proton is moving along the electric field lines, it is gaining energy.
Thus, when a proton moves in the direction of the electric field lines, it is moving from low to high potential and gaining electric potential energy.
Learn more about electric field lines.
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