When does recombination occur? at the two-strand stage, when the sister chromatids have been separated, and after dna replication at the four-strand stage, when each member of a homologous chromosome pair is composed of two sister chromatids, and after dna replication at the two-strand stage, when each member of a homologous chromosome pair is composed of two sister chromatids, and before dna replication at the four-strand stage, when each member of a homologous chromosome pair is composed of two sister chromatids, and before dna replication before dna replication?